Center for Theoretical Study
Jilská 1, 110 00 Praha 1, Czech Rep., phone:+420-221183535, fax: +420-2-22220653, e-mail:

David Storch, Ph.D.

Employment and fellowships


Other professional experience

  • editor of the journal Ecology Letters
  • member of editorial board of the journal Vesmír (The Universe), about 40 popular scientific papers in this and other Czech journals
  • Chair of the council of the Czech Society for Ecology (since 2011)
  • field experience: Caucasus (1989), Siberia, Lake Baikal (1992), South-east Turkey (1996), The Balkans (1987, 1993, 1994, 1995, 1996, 1999, 2000, 2001, 2003, 2005), South Africa (2001, 2005, 2007, 2009, 2012), New Mexico (2003, 2004, 2006, 2009, 2014), Costa Rica (2007), Mexico (2009), Papua New Guinea (2010), Cameroon (2011), Namibia, Botswana & Zimbabwe (2012, 2014)

Research interests

Macroecology, biodiversity, ecological theory, biogeography, community ecology


University courses

  • Animal ecology (Ekologie živočichů; together with Lukáš Kratochvíl)
  • Macroecology (Makroekologie)
  • Ecological theory and community ecology (Ekologická teorie a ekologie společenstev; together with prof. Tomáš Herben)
  • Field course of quantitative ornithology (Terénní cvičení z ornitologie)
  • Macroecology and Biodiversity (Makroekologie a biodiverzita; at University of South Bohemia)


Selected publications


Original papers (see Google Scholar or ResearchGate)

Keil P., Storch D., Jetz W. 2015. On the decline of biodiversity due to area loss. Nature Communications, DOI:10.1038/ncomms9837. [PDF]

Miko L. & Storch D. 2015. Biodiversity conservation under energy limitation: Possible consequences of human productivity appropriation for species richness, ecosystem functioning, and food production. Ecosystem Services 16: 146-149. [PDF]

Okie J.G., Van Horn D.J., Storch D., Barrett J.E, Gooseff M.N., Kopsova L., Takacs-Vesbach C.D. 2015: Niche and metabolic principles explain patterns of diversity and distribution: theory and a case study with soil bacterial communities. Proceedings of the Royal Society B., DOI: 10.1098/rspb.2014.2630. [PDF]

Šímova I., Violle C., Kraft N., Storch D., Svenning J.-C., Boyle B., Donoghue J., Jorgensen P.M., McGill B. J., Morueta-Holme N., Peet R.K., Wiser S.K., Piel W., Regetz J., Shildhauer M., Thiers B. & Enquist B.J. 2015: Shifts in trait means and variances in North American tree assemblages: species richness patterns are loosely related to the functional space. Ecography 38: 649-658 (Editor's choice). [PDF]

Hořák D., Tószögyová A. & Storch D. 2015: Relative food limitation drives geographic clutch size variation in South African passerines: a large-scale test of Ashmole’s seasonality hypothesis. Global Ecology and Biogeography 24: 437-447. [PDF]

Varela S., Lima-Ribeiro M.S., Diniz-Filho J.A.F. & Storch D. 2015: Differential effects of temperature change and human impact on European Late Quaternary mammalian extinctions. Global Change Biology 21: 1475-1481. [PDF]

Marquet P.A., Allen A.P., Brown J.H., Dunne J.A., Enquist B.J., Gillooly J.F., Gowaty P.A., Green J.L., Harte J, Hubbell S.P., O´Dwyer J., Okie J.G., Ostling A., Ritchie M., Storch D. & West G.B. 2014: On theory in ecology. BioScience 64: 701-710. [PDF]

Tropek R., Sedláček O., Beck J., Keil P., Musilová Z., Šímová I. & Storch D. 2014: Comment on "High-resolution global maps of 21st-century forest cover change". Science 344: 981-d, DOI: 10.1126/science.1248753. [PDF]

Ferenc M., Sedláček O., Fuchs R., Dinetti M., Fraissinet M. & Storch D. 2014: Are cities different? Patterns of species richness and beta diversity of urban bird communities and regional species assemblages in Europe. Global Ecology and Biogeography 23: 479-489. [PDF]

Pe’er G., Tsianou M.A., Franz K.W., Matsinos Y.G., Mazaris A.D., Storch D., Kopsová L., Verboom J., Baguette M., Stevens V.M. & Henle K. 2014: Toward better application of minimum area requirements in conservation planning. Biological Conservation 170: 92-102. [PDF]

Macháč A., Storch D. & Wiens J.J. 2013: Ecological causes of decelerating diversification in carnivoran mammals. Evolution 67-8: 2423-2433. [PDF]

Šizling A.L., Kunin W.E. & Storch D. 2013: Taxon invariances, maximum entropy, and the species-area relationship. American Naturalist 181: 288-290. [PDF]

Šímová I., Li Y.M. & Storch D. 2013: Relationship between species richness and productivity in plants: the role of sampling effect, heterogeneity and species pool. Journal of Ecology 101: 161-170. [PDF]

Isaac N.J.B., Storch D. & Carbone C. 2013: The paradox of energy equivalence. Global Ecology and Biogeography 22: 1-5. [PDF]

Storch D., Keil P. & Jetz W. 2012: Universal species-area and endemics-area relationships at continental scales. Nature 488: 78-81. [PDFSupplement]

Macháč A., Zrzavý J., Smrčková J. & Storch D. 2012: Temperature-dependence of evolutionary diversification: differences between two contrasting model taxa support the metabolic theory of ecology. Journal of Evolutionary Biology 25: 2449-2456. [PDF]

Keil P., Schweiger O., Kühn I., Kunin W.E., Kuussaari M., Settele J., Henle K., Brotons L., Peer G., Lengyel S., Moustakas A., Steinicke H. & Storch D. 2012: Patterns of beta diversity in Europe: the role of climate, land-cover and distance across scales. Journal of Biogeography 39: 1473-1486. [PDF]

Šizling A.L., Kunin W.E., Šizlingová E., Reif J. & Storch D. 2011: Between geometry and biology: the problem of universality of the species-area relationship. American Naturalist 178: 602-611. [PDF, Appendix A, AppendixB]

Hořák D., Sedláček O., Tószögyová A., Albrecht T., Ferenc M., Jelínek V. & Storch D. 2011: Geographic variation in avian clutch size and nest predation risk along a productivity gradient in South Africa. Ostrich 82: 175-183. [PDF]

Šímová I., Storch D., Keil P., Boyle B., Phillips O.L. & Enquist B.J. 2011: Global species-energy relationship in forest plots: role of abundance, temperature and species' climatic tolerances. Global Ecology and Biogeography 20: 842-856. [PDF]

Isaac N.J.B., Storch D. & Carbone C. 2011: Taxonomic variation in size-density relationships challenges the notion of energy equivalence. Biology Letters 7: 615-618. [PDF]

Macháč A., Zrzavý J. & Storch D. 2011: Range size heritability in Carnivora is driven by geographic constraints. American Naturalist 177: 767-779. [PDF]

Pautasso M., Böhning-Gaese K., Clergeau P., Cueto V.R., Dinetti M., Fernández-Juricic E., Kaisanlahti-Jokimäki M.-L., Jokimäki J., McKinney M.L., Sodhi N.S., Storch D., Tomialojc L., Weisberg P.J., Woinarski J., Fuller R.A. & Cantarello E. 2011: Global macroecology of bird assemblages in urbanized and semi-natural ecosystems. Global Ecology and Biogeography 20: 426-436. [PDF]

Keil P., Herben T., Rosindell J. & Storch D. 2010: Predictions of Taylor's power law, density dependence and pink noise from a neutrally modeled time series. Journal of Theoretical Biology 265: 78-86. [PDF]

Šizling A.L., Storch D. & Keil P. 2009: Rapoport's rule, species tolerances, and the latitudinal diversity gradient: geometric considerations. Ecology 90: 3575-3586. [PDF]

Harte J., Smith A.B. & Storch D. 2009: Biodiversity scales from plots to biomes with a universal species-area curve. Ecology Letters 12: 789-797. [PDF]

Šizling A.L., Šizlingová E., Storch D., Reif J. & Gaston K.J. 2009: Rarity, commonness and the contribution of individual species to species richness patterns. American Naturalist 174: 82-93. [PDF]

Morlon H., White E.P., Etienne R.S., Green J.L., Ostling A., Alonso D., Enquist B.J., He F., Hurlbert A., Magurran A.E., Maurer B.A., McGill B.J., Olff H., Storch D. & Zillio T. 2009: Taking species abundance distributions beyond individuals. Ecology Letters 12: 488-501. [PDF]

Šizling A.L., Storch D., Šizlingová E., Reif J. & Gaston K.J. 2009: Species-abundance distribution results from a spatial analogy of central limit theorem. Proceedings of the National Academy of Sciences of the U.S.A. 106: 6691-6695. [PDF]

Šizling A.L., Storch D., Reif J. & Gaston K.J. 2009: Invariance in species-abundance distribution. Theoretical Ecology 2: 89-103. [PDF]

Reif J., Storch D. & Šímová I. 2008: The effect of scale-dependent habitat gradients on the structure of bird assemblages in the Czech Republic. Acta Ornithologica 43: 197-206. [PDF]

Storch D. & Šizling A.L. 2008: The concept of taxon invariance in ecology: do diversity patterns vary with changes in taxonomic resolution? Folia Geobotanica 43: 329-344. [PDF]

Nekola J.C., Šizling A.L., Boyer A.G. & Storch D. 2008: Artifactions in the log-transformation of species abundance distributions. Folia Geobotanica 43: 259-268. [PDF]

Reif J., Storch D., Voříšek P., Šťastný K. & Bejček V. 2008: Bird-habitat associations predict population trends in central European forest and farmland birds. Biodiversity and Conservation 17: 3307-3319. [PDF]

Keil P., Dziock F. & Storch D. 2008: Geographical patterns of hoverfly (Diptera, Syrphidae) functional groups in Europe: inconsistency in environmental correlates and latitudinal trends. Ecological Entomology 33: 748-757. [PDF]

Storch D., Šizling A.L., Reif J., Polechová J., Šizlingová E. & Gaston K.J. 2008: The quest for a null model for macroecological patterns: geometry of species distributions at multiple spatial scales. Ecology Letters 11: 771-784. [PDF]

Evans K.L., Newson S.E., Storch D., Greenwood J.J.D. & Gaston K.J. 2008: Spatial scale, abundance and the species-energy relationship in British birds. Journal of Animal Ecology 77: 395-405. [PDF]

Davies R.D., Orme C.D.L., Storch D., Olson V., Thomas G.H., Ross S.G., Benett P.M., Owens I.P.F., Blackburn T.M. & Gaston K.J. 2007: Topography, energy and the global distribution of bird species richness. Proceedings of the Royal Society of London B 274: 1189-1197. [PDF]

Storch D., Davies R.G., Zajíček S., Orme C.D.L., Olson V., Thomas G.H., Ding T.S., Rasmussen P.C., Ridgely R.S., Bennett P.M., Blackburn T.M., Owens I.P.F. & Gaston K.J. 2006: Energy, range dynamics and global species richness patterns: Reconciling mid-domain effects and environmental determinants of avian diversity. Ecology Letters 9: 1308-1320. [PDF]

Reif J., Hořák D., Sedláček O., Riegert J., Pešata M., Hrázský Z., Janeček Š. & Storch D. 2006: Unusual abundance-range size relationship in an Afromontane bird community: the effect of geographical isolation? Journal of Biogeography 33: 1959-1968. [PDF]

Storch D., Evans K. L. & Gaston K. J. 2005: The species-area-energy relationship. Ecology Letters 8: 487-492. [PDF]

Storch D., Marquet P. A. & Gaston K. J. 2005: Untangling an entangled bank. Science 307: 684-686. [PDF]

Storch D. & Gaston K. J. 2004: Untangling ecological complexity on different scales of space and time. Basic and Applied Ecology 5: 389-400. [PDF]

Bonn A., Storch D. & Gaston K. J. 2004: Structure of the species-energy relationship. Proceedings of the Royal Society of London B 271: 1685-1691. [PDF]

Šizling A.L. & Storch D 2004: Power-law species-area relationships and self-similar species distributions within finite areas. Ecology Letters   7: 60-68. [PDF]

Storch D. 2003: Comment on “Global biodiversity, biochemical kinetics and the energetic-equivalence rule”. Science 299: 346b. [PDF]

Storch D., Šizling A. & Gaston K. 2003: Geometry of the species-area relationship in central European birds: testing the mechanism. Journal of Animal Ecology 72: 509-519. [PDF]

Storch D., Konvička M., Beneš J., Martinková J. & Gaston K.J. 2003: Distribution patterns in butterflies and birds of the Czech Republic: separating effects of habitat and geographic position. Journal of Biogeography 30: 1195-1205. [PDF]

Storch D., Gaston K.J. & Cepák J. 2002: Pink landscapes: 1/f spectra of spatial environmental variability and bird community composition. Proceedings of the Royal Society of London B 269: 1791-1796. [PDF]

Storch D. & Šizling A.L. 2002: Patterns in commoness and rarity in central European birds: Reliability of the core-satellite hypothesis within a large scale. Ecography 25: 405-416. [PDF]

Storch D. & Frynta D. 2000: Evolution of habitat selection: Stochastic acquisition of cognitive clues? Evolutionary Ecology 13: 591-600. [PDF]

Storch D.2000: Rapoport effect and speciation/extinction rates in the tropics. Trends in Ecology and Evolution 15: 514.

Storch D. & Kotecký V. 1999: Structure of bird communities in the Czech Republic: The effect of area, census technique and habitat type. Folia zoologica 48 (4): 265-277.

Storch D. & Musil P. 1999: Waterbird habitat preferences and their inter-annual changes related to population dynamics. Vogelwelt 120 (Suppl.): 257-260.

Storch D. 1998: Densities and territory sizes of birds in two different  lowland communities in eastern Bohemia. Folia zoologica 47 (3): 181-188.



Zrzavý J., Storch D. & Mihulka S. 2009: Evolution: Ein Lese-Lehrbuch. Spektrum Akademischer Verlag, Heidelberg.

Storch D., Marquet P.A. & Brown J.H. (eds.) 2007: Scaling biodiversity. Cambridge University Press, Cambridge.

Zrzavý J., Storch D. & Mihulka S. 2004: Jak se dělá evoluce: Od sobeckých genů k rozmanitosti života. [Evolution: From selfish genes to the diversity of life] Paseka, Praha.

Sádlo J., Storch D. 2000: Biologie krajiny: Biotopy České republiky. [Biology of Landscape: Habitats of the Czech Republic] Vesmír, Praha.

Storch D., Mihulka S. 2000: Úvod do současné ekologie. [An introduction to contemporary ecology] Portál, Praha.

Storch D., Mihulka S. 1997: Ekologie. [Ecology] IDM MŠMT ČR, Praha. [the first, shorter version of Storch & Mihulka 2000]


Reviews, book chapters and major popular scientific articles

Storch D. 2016: Záhada mizejícího vrabčáka a jiné proměny české přírody. Lidové Noviny - Orientace, 30. ledna 2016, str. 24-25. [PDF]

Storch D. 2015: Fylogeneze ptáků: Vyřešeno? Vesmír 94: 556-563. [PDF; příloha s fylogenetickým stromem PDF; dodatek PDF]

Storch D. 2015: Máme vizi ochrany přírody? Fórum ochrany přírody 01/2015: 12-14 [PDF]

Storch D., Keil P. & Kunin W.E. 2014: Scaling communities and biodiversity. In. Henle K., Potts S.G., Kunin W.E., Matsinos Y.G., Similä J., Pantis J.D., Grobelnik V., Penev L. & Settele J. (eds.), Scaling in ecology and biodiversity conservation. Pensoft Publishers, Sofia, pp. 66-77.  [PDF]

Storch D. 2014: Věda z Marsu: Úskalí ekologie podle Google Earth aneb Jak je důležité neplést si sóju s pralesem. Respekt 26, 36.-29. červen 2014, str. 70-71.  [PDF]

Storch D. 2014: Co dělat? Vesmír 93: 241.

Storch D. 2014: Hledání ztraceného ráje. Lidové Noviny - Orientace, 11. ledna 2014, str. 21-22. [PDF]

Storch D. 2013: Sovy nemusejí být, čím se zdají [projev při převzetí profesorského dekretu]. Lidové Noviny - Orientace, 29. června 2013, str. 30. [PDF]

Storch D. 2013: Ochrana přírody jako vědecký problém. Vesmír 92: 113-115. [PDF]

Storch D. 2012: Hlad po fosforu. Respekt 45, 5.-11. listopad 2012, str. 63-65. [PDF]

Storch D. 2012: Svátek přehnojené planety: Přírodu má smysl chránit - ale z úplně jiných důvodů a úplně jinak, než si lidé myslí. Lidové Noviny - Orientace, 21.-22. dubna 2012, str. 21-22. [PDF]

Storch D. 2012: Věda, ekologie, ideologie a myšlení. In: Hlaváček P. (ed.), Intelektuál ve veřejném prostoru: Vzdělanost, věda, politika. Academia, Praha, pp. 343-351. [PDF]

Storch D. 2012: Biodiversity and its energetic and thermal controls. In: Sibly R.M., Brown J.H. & Kodric-Brown A. (eds.), Metabolic Ecology: A Scaling Approach. Wiley-Blackwell, pp. 120-131. [PDF]

Storch D. 2011: Žijeme v době šestého masového vymírání? Odhady současných změn biodiverzity po deseti letech. Vesmír 90: 568-572. [PDF; příloha PDF]

Storch D. 2011: Teorie ideálního lesa. Vesmír 90: 351-356. [PDF]

Storch D. 2010: Metabolismus a velikost těla v novém tisíciletí: Je tříčtvrtinový koeficient univerzální? Vesmír 89: 536-538. [PDF]

Storch D. 2010: Dvě vědy: Matematická přírodověda, taxonomie a přirozený svět. In: Velický B., Trlifajová K., Kouba P. et al., Spor o přirozený svět. Filosofia, Praha, pp. 273-284. [PDF]

Storch D. & Šizling A.L. 2009: O převaze vzácnosti v přírodě: Dobře ukrytá informace o struktuře krajiny. Vesmír 88: 784-787. [PDF]

Storch D. 2009: Čtení Darwina: Původ druhů po 150 letech. Vesmír 88: 444-450. [PDF]

Polechová J. & Storch D. 2008: Ecological niche. In: Jorgensen S.E. & Fath B. (eds.), Encyclopedia of Ecology. Elsevier, Oxford, pp. 1088-1097. [PDF]

Storch D. 2008: Katastrofy v přírodě: statistika, dynamika a teorie. In: Pokorný P. & Bárta M. (eds.), Něco překrásného se končí: Kolapsy v přírodě a společnosti. Dokořán, Praha, pp. 31-36. [PDF]

Storch D., Šizling A.L. & Gaston, K.J. 2007: Scaling species richness and distribution: Uniting the species-area and species-energy relationships. In: Storch D., Marquet P.A. & Brown J.H. (eds.), Scaling biodiversity. Cambridge University Press, Cambridge, pp. 300-322. [PDF]

Šizling A.L. & Storch D. 2007: Geometry of species distributions: Random clustering and scale invariance. In: Storch D., Marquet P.A. & Brown J.H. (eds.), Scaling biodiversity. Cambridge University Press, Cambridge, pp. 77-100. [PDF]

Storch D. 2006: Jaký je vlastně les: Obraz lesa v přírodních vědách. Dějiny a současnost 11: 34-36. [PDF]

Storch D. 2006: Pochybnosti o stavu přírody aneb proč ji vlastně chránit. Ekolist 3: 12-15. [PDF]

Storch D. 2004: Metabolická teorie biologie aneb Nová teorie všeho (živého)? Vesmír 83, 508-516. [PDF]

Storch D. and Reif J. 2003: Macroecology of birds: what have we learned from the large-scale censuses. Sylvia 38, 1-18. (in Czech, summary in English) [PDF]

Storch D. 2002: Krajina není. In: Hájek P. (editor) 2002: Krajina zevnitř. Malá skála, Praha.

Storch D. 2001: Biodiversity, Chapter in Encyclopedia of Life support Systems, UNESCO.

Storch D. 2001: Jak rychle vymírají druhy: Potíže s odhady globální biodiverzity a jejích změn. Vesmír 80: 573-575. [PDF]

Storch D. 2000: O vzniku druhů přirozeným výběrem: Role prostředí v evoluci biodiverzity. Vesmír 79: 550-552. [PDF]

Storch D. 2000: Přežívání populací v ostrůvkovitém prostředí: Co jsou metapopulace a jak fungují. Vesmír 79: 143-145. [PDF]

Storch D. 1999: Proč je malých zvířat víc než velkých. Vesmír 78: 377-383. [PDF]

Storch D. 1998: Proč žije v tropech tolik druhů organismů. Vesmír 77: 677-678. [PDF]

Storch D. 1995: Interspecific competition in birds. Biol.listy  60: 259-273. (in Czech, summary in English)


Invited lectures


  • Storch D.: Do we understand spatial diversity patterns? The origin of universal laws in macroecology (58th Annual Symposium of IAVS: Understanding broad-scale vegetation patterns, Brno, Czech Republic, 19-24 July 2015)
  • Storch D.: On the origin of biodiversity patterns: Towards equilibrium diversity theory (iDiv, German Centre for Integrative Biodiversity Research, Leipzig, 24 June 2015)
  • Storch D.: Are there general patterns in ecology? Invariance and universality in macroecology (Santa Fe Institute, USA, 29 August 2014)
  • Storch D.: Do we understand diversity? On the origin of universal macroecological patterns (Yale Institute for Biospheric Studies, Yale University, USA, January 2013)
  • Storch D.: Species extinction due to habitat loss: On a proper use of species-area and endemics-area relationships (Gordon Research Conference on Metabolic Basis Of Ecology, University of New England in Biddeford, ME, USA, 22-27 July 2012)
  • Storch D.: Invariances in macroecology: Are species-abundance, species-area, and species-energy relationships universal? (12th congress of European Ecological Federation, Ávilla, Spain, 25-29 September 2011)
  • Storch D.: Are there general laws of landscape structure? Scaling spatial distributions, beta diversity, and the species-area relationship. (International Conference in Landscape Ecology, Brno, Czech Republic, 3-6 September 2010)
  • Storch D.: Thermal and energetic controls on biodiversity (Gordon Research Conference on Metabolic Basis Of Ecology, University of New England in Biddeford, ME, USA, 18-23 July 2010)
  • Storch D.: Metabolic theory of biodiversity? (British Ecological Society annual meeting, University of Hertfordshire, 7-10 September, 2009)
  • Storch D.: The relationship between energy availability and species diversity: role of abundance, range dynamics and speciation (University of Arizona, Tucson, USA, February 25, 2008)
  • Storch D.: Abundance, distribution and diversity: The quest for a null model of biodiversity patterns (University of California, San Diego, USA, October 13, 2006)
  • Storch D.: Geometry of species distributions: null models of biodiversity patterns (Santa Fe Institute, USA, September 15-19, 2006).
  • Storch D.: Macroecological patterns (Workshop "Biodiversity Patterns, Ecological Modelling & Statistics", Moulis, France, August 23-September 7, 2006)
  • Storch D. & Šizling A.L.: Geometry of species distributions: null models of biodiversity patterns (International conference "Macroecological Tools for Global Change Research", Potsdam, Germany, August 21-23, 2006).
  • Storch D.: Linking biodiversity patterns to species´ spatial distributions: scaling patterns in continental avifaunas derived from the structure and dynamics of species´ ranges (24th International Ornithological Congress, Hamburg, Germany, August 13-19, 2006).
  • Storch D. & Šizling A.L.: From species to communities: the scaling of biodiversity (Annual meeting of Ecological Society of America and IX International Congress of Ecology, INTECOL, Montreal, Canada, August 7-12, 2005)
  • Storch D.: Major ultimate determinants of (avian) species richness patterns (Department of Biology, University of New Mexico, USA, March 2003)
  • Storch D.: Patterns in animal abundance and distribution: reliability of null models (Santa Fe Institute, USA, February 2003)
  • Storch, D.: Macroecological patterns in distribution of organisms: Complex patterns with simple explanations and simple patterns with complex explanations (Joint workshop of Santa Fe Institute for the Sciences of Complexity and Max Planck Institute for Mathematics in the Sciences, Leipzig, Germany, 2001)
  • Storch D.: Habitat selection: Ecology and evolution (Department of Zoology, University of Oxford, UK, November 2000)
  Picture of David Storch

Scaling biodiversity

Jak se dela evoluce

Biologie krajiny

Uvod do soucasne ekologie

Evolution: Ein Lese-Lehrbuch